How does mail routing work?

Mail could furthermore be routed through the network, traversing any number of intermediate nodes before arriving at its destination. Initially, this had to be done by specifying the complete path, with a list of intermediate host names separated by bangs. For example, if machine barbox is not connected to the local machine, but it is known that barbox is connected to machine foovax which does communicate with foovax, the appropriate address to send mail to would be foovax!barbox!you.